The second cercopithecoid subfamily – confusingly known as the Cercopithecinae – diversified at around the same time as the Colobinae. They split into two tribes. One includes some of Africa’s most beautiful species including the guenons, patas monkey, green monkeys and vervets. The other includes baboons, geladas, mangabeys, mandrills and macaques. This second group in particular has a rich history of hybridisation.

Images by Hans Hillewaert (De Brazza’s monkey, far left) Muhammad Mahdi Karim (crab-eating macaque, left), me (hamadryaz baboon, right) Malene Thyssen (mandrill, far right)